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Dr. Markus Koch

Wissenschaftlicher Mitarbeiter

Markus Koch


  • Biodiversity and evolutionary morphology of soil arthropods
  • Origin and early evolution of insects (Hexapoda)
  • Systematics and integrative taxonomy of apterygote hexapods (focus: Diplura, Zygentoma)
  • Phylogeny of myriapods
  • Morphogenesis of arthropods (focus: organogenesis, evolution of the mesoderm)
  • Anatomical ontologies and information technology (BIOfid, Specialized Information Service for Biodiversity Research)


Bachelor-Programme Biology

  • BIO-02 Morphologie und Evolution der Tiere
  • BIO-07 Ökologie mit Bestimmungsübungen
  • WBIO-C-04 Biodiversität und Phylogenie der Metazoa

International Master-Programme "Organismic Biology, Evolutionary Biology and Palaeobiology" (OEP Biology)

  • OEP-M1 Biodiversity & Evolution
  • OEP-M2 Fundamentals of Evolutionary Biology
  • OEP-A07 Histology, tomography, and computer-aided 3D-reconstruction of animal anatomy
  • OEP-A08 Morphological character analysis in phylogenetics
  • OEP-A11 Practical course on electron microscopy

Akademische Laufbahn

since Oct 2020
Senior Researcher, Institute of Evolutionary Biology and Ecology, University of Bonn
 Sept-Dez 2018 Visiting Professor, University of Córdoba (Spain)


Research Associate

  • Senckenberg Society for Nature Research, Leibniz Institution for Biodiversity and Earth System Research
  • Institute of Evolutionary Biology and Ecology, University of Bonn
2014-2017 Akademischer Rat auf Zeit, Institute of Evolutionary Biology and Ecology, University of Bonn


Interim Professor for Systematics and Biodiversity of Insects, Biocentre Grindel and Zoological Museum, University of Hamburg

2009-2011 Akademischer Rat auf Zeit, Institute of Evolutionary Biology and Ecology, University of Bonn

Assistant Professor, Institute of Biology, Dept. Animal Systematics and Evolution, Freie Universität Berlin

2000-2002 Postdoc, Faculty of Biology, Dept. Zoomorphology & Systematics, University of Bielefeld
1996-2000 Ph.D. student (Zoological Systematics), University of Hamburg
1989-1995 Diploma student (Biology), University of Hamburg

Ausgewählte Publikationen

Full list of publications on ORCID.

Information technology

  • González-Montaña, L.A. & Koch, M. (2021) Lepidoptera Anatomy Ontology (LepAO) (Version v2021-11-20). GitHub repository, doi: 10.5281/zenodo.4780615.
  • Driller, C., Koch, M., Abrami, G., Hemati, W., Lücking, A., Mehler, A., Pachzelt, A. & Kasperek, G. (2020) Fast and easy access to Central European biodiversity data with BIOfid. Biodiversity Information Science and Standards 4: e59157.
  • Koch, M., Driller, C., Schmidt, M., Hörnschemeyer, T., Weiland, C., Miko, I., Yoder, M. & Hickler, T. (2018) Current progress in the development of taxonomic and anatomical ontologies within the scope of BIOfid. – Biodiversity Information Science and Standards 2: e25585.
  • Driller, C., Koch, M., Schmidt, M., Weiland, C., Hörnschemeyer, T., Hickler, T., Abrami, G., Ahmed, S., Gleim, R., Hemati, W., Uslu, T., Mehler, A., Pachzelt, A., Rexhepi, J., Risse, T., Schuster, J., Kasperek, G. & Hausinger, A. (2018) Workflow and current achievements of BIOfid, an information service mobilizing biodiversity data from literature sources. – Biodiversity Information Science and Standards 2: e25876.
  • Koch, M., Kasperek, G., Hörnschemeyer, T., Mehler, A., Weiland, C. & Hausinger, A. (2017): Setup of BIOfid, a new Specialised Information Service for Biodiversity Research. – Biodiversity Information Science and Standards 1: e19803.

Evolution of Myriapods

  • Edgecombe, G.D., Strullu-Derrien, C., Hetherington, A.J., Goral, T. & Koch, M. (2020) Aquatic stem group myriapods close a gap between molecular divergence dates and the terrestrial fossil record. PNAS 117 (16): 8966-8972.
  • Moritz, L. & Koch, M. (2020) No Tömösváry organ in flat backed millipedes (Diplopoda: Polydesmida). ZooKeys 930 (Special Issue "Proceedings of the 18th International Congress of Myriapodology, 25‒31 August 2019, Budapest, Hungary"): 103-115.
  • Moritz, L., Wesener, T. & Koch, M. (2018): An apparently non-swinging tentorium in the Diplopoda (Myriapoda): Comparative morphology of the tentorial complex in giant pill-millipedes (Sphaerotheriida). - Zookeys 741: 77-91.
  • Koch, M. (2015): Diplopoda – General Morphology. – In: A. Minelli (ed.), Treatise on Zoology – Anatomy, Taxonomy, Biology. The Myriapoda, Volume 2, Chapter 2, pp. 7-67. Brill, Leiden.
  • Koch, M. (2015): Diplopoda – Skeletomuscular System and Locomotion. – In: A. Minelli (ed.), Treatise on Zoology – Anatomy, Taxonomy, Biology. The Myriapoda, Volume 2, Chapter 4, pp. 101-107. Brill, Leiden.
  • Koch, M., Schultz, J. & Edgecombe, G.D. (2015): Tentorial mobility in centipedes (Chilopoda) revisited: 3D reconstruction of the mandibulo-tentorial musculature of Geophilomorpha. – Zookeys 510: 243-267.
  • Koch, M. & Edgecombe, G.D. (2012): The preoral chamber in geophilomorph centipedes: comparative morphology, phylogeny and the evolution of centipede feeding structures. – Zoological Journal of the Linnean Society 165: 1-62.
  • Koch, M., Edgecombe, G.D. & Shelley, R. M. (2010): Anatomy of Ectonocryptoides (Scolopocryptopidae: Ectonocryptopinae) and the phylogeny of blind Scolopendromorpha (Chilopoda). – International Journal of Myriapodology 3: 51-81.
  • Koch, M., Pärschke, S. & Edgecombe, G.D. (2009): Phylogenetic implications of gizzard morphology in scolopendromorph centipedes (Chilopoda). – Zoologica Scripta 38: 269–288.

Evolution of the Mesoderm

  • Koch, M., Quast, B. & Bartolomaeus, T. (2014): Coeloms and nephridia in annelids and arthropods. – In: Waegele, W. & Bartolomaeus, T. (eds.), Deep Metazoan Phylogeny: The Backbone of the Tree of Life. New insights from analyses of molecules, morphology, and theory of data analysis. Chapter 11, pp. 173-284. DeGruyter.

Evolution of Insects (Hexapoda)

  • Koch, M. (2016): Current views on the early evolution of the insect head revisited. – Zitteliana 88: 31.
  • Blanke, A., Koch, M., Wipfler, B., Wilde, F. & Misof, B. (2014): Head morphology of Tricholepidion gertschi indicates monophyletic Zygentoma. – Frontiers in Zoology, 11:16.
  • Chen, W.J., Koch, M., Mallat, J.M. & Luan, Y.X. (2014): Comparative analysis of mitochondrial genomes in Diplura (Hexapoda, Arthropoda): taxon sampling is crucial for phylogenetic inferences. - Genome Biology and Evolution, 6(1):105–120.
  • Hörnschemeyer, T., Haug, J., Bethoux, O., Beutel, R.G., Charbonier, S., Hegna, T.H., Koch, M., Rust, J., Wedmann, S., Bradler, S. & Willmann, R. (2013): Is Strudiella a Devonian insect? – Nature 494: E3-E4 (DOI 10.1038/nature11887).
  • Blanke, A., Wipfler, B., Letsch, H., Koch, M., Beckmann, F., Beutel, R.G. & Misof, B. (2012): Revival of Palaeoptera – head characters support a monophyletic origin of Odonata and Ephemeroptera (Insecta). – Cladistics 28: 560-581.
  • Koch, M. (2009): Biodiversity of the two-pronged bristletails (Diplura) in Western Australia as revealed from recent mining projects. – Enviromental Protection Authority Report (Government of Western Australia), Tropicana Gold Project, Supplementary Study 3k.


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